Grug （瓜哥）- The protagonist, a strong and somewhat stubborn primitive father who is deeply attached to tradition and family. He is resistant to new ideas and unwilling to change his ways.

Ugga （乌娜）- Grug's wife, a gentle mother who is open to new ideas and always tries to find solutions to Grug's stubbornness. She often attempts to find a way to make Grug see the value of change.

Eep （小伊）- Grug and Ugga's daughter, a curious and somewhat rebellious young girl who longs to explore the world and break free from her family's traditional bonds. She is adventurous and eager to learn about the outside world.

Guy （盖）- A clever wanderer who brings new information and survival skills to the tribe. He tries to persuade Grug and the others to accept his new ideas, but he often faces rejection.

The "Other" Primitives- A group of primitives who embrace Guy's thoughts and begin to change their lifestyle to adapt to the new environment. They serve as a contrast to Grug's family.

These characters form the heart of the "The Croods" universe, each with their unique traits and roles in the story.